The Control of Microbial Growth

 

The Terminology of Microbial Control

 

1       The control of microbial growth can prevent infections and food spoilage.

2       Sterilization is the process of destroying all microbial life on an object.

3       Commercial sterilization is heat treatment of canned foods to destroy C. botulinum endospores.

a.     Resistant thermophilic endospores don’t germinate at food storage temperatures – the problem comes at temperatures above 45° C.

4       Disinfection is the process of reducing or inhibiting microbial growth on a nonliving surface.

5       Antisepsis is the process of reducing or inhibiting microorganisms on a living tissue.

6       The suffix -cide means to kill; the suffix -stat means to inhibit

7       Sepsis is bacterial contamination.

 

The Rate of Microbial Death

 

1       Bacterial populations subjected to heat or antimicrobial chemicals usually die at a constant rate.

2       Such a death curve, when plotted logarithmically, shows this constant death rate as a straight line.

3       The time it takes to kill a microbial population is proportional to the number of microbes.

4       Microbial species and life cycles phases (endospores) have different susceptibilities to physical and chemical controls.

5       Organic matter may interfere with heat treatments and chemical control agents.

6       Longer exposure to lower heat can produce the same effect as shorter time at higher heat.

 

Actions of Microbial Control Agents

 

Alterations of Membrane Permeability

 

1       The susceptibility of the plasma membrane is due to its lipid and protein components.

2       Certain chemical control agents damage the plasma membrane by altering its permeability.

 

Damage to Proteins and Nucleic Acids

 

1       Some microbial control agents damage cellular proteins by breaking hydrogen bonds and covalent bonds.

2       Other agents interfere with DNA and RNA replication and protein synthesis.

 

Physical Methods of Microbial Control

 

Heat

 

1       Heat is frequently used to eliminate microorganisms.

2       Most heat kills microbes by denaturing enzymes.

a.     Heat works better at low pH.

3       Thermal death point (TDP) is the lowest temperature at which all the microbes in a liquid culture will be killed in 10 minutes.

4       Thermal death time (TDT) is the length of time required to kill all bacteria in a liquid culture at a given temperature.

5       Decimal reduction time (DRT) is the length of time in which 90% of a bacterial population will be killed at a given temperature.

6       Boiling (100ş C) kills many vegetative cells and viruses within 10 minutes.

7       Autoclaving (steam under pressure) is the most effective method of moist heat sterilization. The steam must directly contact the material to be sterilized.

a.     Fifteen psi, 121 ° C for fifteen minutes kills all vegetative bacteria and endospores.

8       In HTST pasteurization, a high temperature is used for a short time (72ş C for 15 seconds) to destroy pathogens without altering the flavor of the food. Ultra-high-temperature (UHT) treatment (140ş C for 3 seconds) is used for dairy products.

a.     Classic pasteurization was done at 63° C for 30 minutes

9       Methods of dry heat sterilization include direct flaming, incineration, and hot -air sterilization (170° C for two hours). Dry heat kills by oxidation.

10    Different methods that produce the same effect (reduction in microbial growth) are called equivalent treatments.

 

Filtration

 

1       Filtration is the passage of a liquid or gas through a filter with pores small enough to remain microbes.

2       Microbes can be removed from air by high -efficiency particulate air filters.

a.     HEPA – 0.3 um pore size

b.     Decreases numbers

3       Membrane filters composed of nitrocellulose or cellulose acetate are commonly used to filter out bacteria, viruses, and even large proteins.

a.     Range from 0.22 um – 0.45 um for bacteria but sometimes mycoplasma or spirochetes can pass through.

b.     Sizes can go as low as 0.01 um to filter viruses and proteins.

 

Low Temperatures 

 

1       The effectiveness of low temperatures depends on the particular microorganism and the intensity of the application.

2       Most microorganisms do not reproduce at ordinary refrigerator temperatures (0-7ş C).

3       Many microbes survive (but do not grow) at subzero temperatures used to store foods.

 

High Pressure

 

1       High pressure denatures proteins in vegtetative cells. 

2       Endospores aren’t usually harmed, but can be allowed to grow out and the vegetative form killed with high pressure.

3       Used in Japan and the U.S. to preserve fruit juices – preserves the flavor, color, and nutrient values.

 

Desiccation 

 

1       In the absence of water, microorganisms can grow but cannot remain viable.

2       Viruses and endospores can resist desiccation.

a.     Mycobacterium tuberculosis is fairly resistant to desiccation but the gonorrhea bacterium is very sensitive.

3       Environmental conditions effect efficiency of desiccation – pus, feces, and mucus are protective

 

Osmotic Pressure 

 

1       Microorganisms in high concentrations of salts and sugars undergo plasmolysis.

2       Molds and yeasts are more capable than bacteria of growing in materials with low moisture or high osmotic pressure.

 

Radiation

1       The effects of radiation depend on its wavelength, intensity and duration.

2       Ionizing radiation (gamma rays, X-rays and high-energy electron beams) has a high degree of penetration and exerts its effect primarily by ionizing water and forming highly reactive hydroxyl radicals.

a.     Destroys DNA.

3       Non-ionizing radiation causes damage to DNA.

a.     Ultraviolet causes formation of thymine dimers.

b.     Best wavelength is 260 nm.

4       Microwaves heat water but it is possible to isolate vegetative forms from microwave ovens – probably protected by uneven heating in solid foods

 

Chemical Methods of Microbial Control

 

1       Chemical agents are used on living tissue (as antiseptics) and on inanimate objects (as disinfectants).

2       Few chemical agents achieve sterility.

 

Principles of Effective Disinfection

 

1       Properties of the disinfectant – how it works determines what it will be effective against.

2       Concentration of the disinfectant - requires proper hydration.

3       The presence of organic matter, which can act as a buffer

4       The degree of contact with microorganisms – if the surface needs cleaning or is porous microorganisms can escape contact with the disinfectant.

5       Temperature should also be considered, since increased temperatures usually enhance the efficacy of disinfectants.

a.     Time left in contact is also an important consideration.

 

Evaluating Disinfectants

 

1       In the use-dilution test, bacterial (S. choleraesuis, S. aureus, and P. aeruginosa) survival in the manufacturer’s recommended dilution of a disinfectant is determined.

a.     Viruses, endospore-forming bacteria, mycobacteria, and fungi can also be used in the use-dilution test.

2       In the disk-diffusion method, a disk of filter paper is soaked with a chemical and placed on an inoculated agar plate; a clear zone of inhibition indicates effectiveness.

3       Phenol coefficient – compares activity of phenol.

 

Types of Disinfectants

 

Phenol and Phenolics

1       Phenolics are derivatives of phenol that have been altered to reduce irritating qualities or increase antimicrobial activity when combined with detergents.

2       Phenolics exert their action by injuring plasma membranes.

a.     Can also denature proteins – enzyme inactivation

3       Qualities:

a.     Not inactivated by organic compounds

b.     Stable for long periods

c.     Persist for long periods after application

d.     Good for disinfecting things like pus, saliva, and feces

4       Cresol is a phenolic that is derived from coal tar.

5       O-phenylphenol is the main ingredient in Lysol.

 

 

Bisphenols

 

1       Bisphenols such as triclosan (over the counter) and hexachlorophene (prescription) are widely used in household products.

2       Hexachlorphene is used in pHisoHex

a.     Particularly good against gram-positive strep and staph (cause skin infections in infants)

b.     Can cause neurological damage if exposure is high (bathing several times per day)

3       Triclosan is used in soaps, toothpaste, and incorporated into plastic kitchenware.

a.     Broad spectrum, especially against gram-positive bacteria and fungi

b.     Resistant strains of bacteria have arisen and triclosan may contribute to antibiotic resistance as well

 

Biguanides

 

1       Chlorhexidine

a.     Binds well to skin and mucus membranes

b.     Low toxicity

c.     Surgical hand scrubs and pre-op skin prep

d.     Can damage eyes

e.     Not sporocidal

 

Halogens

 

1       Some halogens (iodine and chlorine) are used alone or as components of inorganic or organic solutions.

2       Iodine may combine with certain amino acids to inactivate enzymes and other cellular proteins.

3       Iodine is available in a tincture (in solution with alcohol) or an iodophor (combined with an organic molecule).

a.     Betadine, Isodine (povidone-iodines)

b.     Povidone is surface active – improves wetting actions.

4       The germicidal action of chlorine is based on the formation of hypochlorous acid (HOCl) when chlorine is added to water.

a.     Good oxidizing agent.

b.     Chlorine is used as a disinfectant in gaseous form (Cl2 ) or in the form of a compound, such as calcium hypochlorite, sodium hypochlorite (NaOCl, Clorox), sodium dichloroisocyanurate, and chloramines.

c.     Used to disinfect drinking water and swimming pools

d.     2 drops Clorox per liter (4 if cloudy) and let sit 30 minutes

 

Alcohols 

1       Alcohols exert their action by denaturing proteins and dissolving lipids.

a.     Not good for wound disinfection because proteins coagulate and form a protective coat around bacteria.

2       In tinctures, they enhance the effectiveness of other antimicrobial chemicals.

3       Aqueous ethanol (60-95%) and isopropanol are used as disinfectants.

a.     Denaturation requires H2O, which is why aqueous preparations are better than pure.

b.     70% is best ethanol concentration.

 

Heavy Metals and Their Compounds

 

1       Silver, mercury, copper, and zinc are used as germicidals.

2       They exert their antimicrobial action through oligodynamic action.  When heavy metal ions combine with sulfhydryl (—SH) groups, proteins are denatured.

 

Surface -Active Agents

 

1       Surface-active agents decrease the surface tension among molecules of a liquid; soaps and detergents are examples.

2       Soaps have limited germicidal action but assist in the removal of microorganisms through scrubbing.

a.     Emulsify fats

3       Acid-anionic detergents are used to clean dairy equipment.

a.     Interact with plasma membrane and may cause damage to enzymes.

 

Quanternary Ammonium Compounds (Quats)

 

1       Quats are cationic detergents attached to NH4+.

2       By disrupting plasma membranes, they allow cytoplasmic constituents to leak out of the cell.

a.     Also denature proteins (inhibit enzymes) and are surface active.

3       Quats are most effective against gram-positive bacteria.

a.     Also good against fungi, amoeba, and enveloped viruses but pseudomonads can grow in them.

4       Inactivated by anions, soaps, detergents, and organic material.

 

Chemical Food Preservatives

 

1       SO2, sorbic acid, benzoic acid, and propionic acid inhibit fungal metabolism and are used as food preservatives.

2       Good to prevent mold growth in acidic foods (molds grow well at low pH)

3       Nitrate and nitrite salts prevent germination of Clostridium botulism endospores in meats and preserve red color.

a.     Nitrites combine with amino acids to produce nitrosamines, which may be carcinogenic.

b.     Nitrosamines are produced from other things in the body and may not be as big a deal as once thought.

 

Antibiotics

 

1       Nisin and natamycin are antibiotics used to preserve foods, especially cheese.

2       Not used for treatment of disease.

 

Aldehydes

 

1       Aldehydes such as formaldehydes and glutaraldehyde exert their antimicrobial effect by inactivating proteins.

2       They are among the most effective chemical disinfectants.

 

Gaseous Chemosterilizers

 

1       Ethylene oxide is the gas most frequently used for sterilization.

2       It penetrates most materials and kills all microorganisms by protein denaturation.

 

Peroxygens  (Oxidizing Agents)

 

1       Ozone peroxide, and peracetic acid are used as antimicrobial agents.

2       They exert their effect by oxidizing molecules inside the cell.

 

Microbial Characteristics And Microbial Control

 

Conditions that Influence Microbial Control

 

1       Temperature

2       Type of Microorganism

a.     Gram-negative bacteria are generally more resistant than gram-positive bacteria to disinfectants and antiseptics.

i)       Pseudomonas and Burkholderia have porins in the cell wall with characteristics that make them resistant to a lot of chemicals and antibiotics.  They can actually grow in some disinfectants and antiseptics due to this resistance combined with the ability to metabolize some unusual molecules.

b.     Mycobacteria, endospores, and protozoan cysts and oocysts are very resistant to disinfectants and antiseptics.

c.     Nonenveloped viruses are generally more resistant than enveloped viruses to disinfectants and antiseptics.

i)       Lipid envelope is a target for chemical injury

d.     Prions resist disinfection and autoclaving - an hour soak in sodium hydroxide followed by an hour of autoclaving at 136 degrees is "fairly effective".

3       Physiological State

a.     Actively growing more susceptible.

b.     Endospores least susceptible.

4       Environment

a.     Organic material inhibits chemicals by binding them and can insulate microorganisms against heat.